TYPE:COLOMBIA.Santander, municipio El Carmen de Chucurí, vereda La Pitala, finca Billete de Oro, cerca de la quebrada La Pitala, 560 m, 6°37'8.3''N, 73°36'9.8''O, 15.II.2019, fl, J. Quiroga-Nova 01 (holotype: JBB!, isotypes: COL!, HUA!, UIS!).Caryodaphnopsis carmensis resembles C. fosteri van der Werff and C. fieldii Aymard & G. A. Romero, but can be differentiated from these species by the golden dense indu-ment on internodes, apical buds, petioles and leaf blades on the adaxial surface, leaf blade oblong to oblong-elliptic, the base subcordate, with 3-5 pairs of secondary veins and with 14-17 lateral veins, prominent and branching toward margin on the adaxial surface. Also, this new species is simi-lar to C. tomentosa van der Werff, but differs from it by the leaf blade oblong to oblong-elliptic, the base subcordate, 3-5 pairs of secondary nerves, and inflorescences single pa-nicles, not or little branched from the base.Trees 20-40 m tall, to 1.5 m in diameter at breast height (DBH), with little buttresses, bark fissured, flaking off in small plates; twigs rectangular in young parts, sub-quadrangular when mature; internodes 3.7-7.6 cm long; dis-tal internodes, terminal buds, petioles and inflorescences covered by a dense golden-brown indumentum, composed by subulate trichomes, 0.3-0.6 mm long; leaves opposite; leaf blade 12.5-23 × 8-12.5 cm, oblong, oblong-elliptic to obovate, thin coriaceous, base rounded to subcordate, apex acute, acuminate, margin entire, glabrous adaxia-lly, with an indument like that of the internodes on veins abaxially; trinerved; midrib and secondary veins impres-sed on the upper surface, elevated on the lower surface; secondary veins 3-5 pairs on each side, alternate, arched and evanescent near margin; lateral veins 14-17, branching towards margin, arching upward, loop-connected and eva-nescent near margin, elevated on the lower surface; petiole 12-19.5 × 2-4 mm, adaxially caniculate to the apex, with an indument like that on the internodes; inflorescence a single panicle, not or little branched from the base, axillary, 1.9-4.8 cm long, paired on distal nodes, sessile or pedunculate, central rachis with 3-5 nodes of ramification, with opposite or alternate branchlets; basal paracladia 16-40 mm long; terminal branches 1-3 flowered; bracts and bracteoles not seen; flowers pedicellate, 1.1-1.3 mm long in bud, cream-green; pedicels 3.8-6.1 mm long, tomentose; tepals 6, in two strongly unequal whorls (persisting in the early stages of the fruit); the outer 1.2-1.4 × 0.8-1.2 mm, wide-ovate, to-mentose externally, glabrous internally; the inner 6.1-6.5 × 2-3.4 mm, ovate, chartaceous, pubescent on both surfaces, trichomes subulate. 0.4-0.9 mm long; stamens 12, in four whorls, each whorl with 3 stamens, three fertile whorls (9 fertile stamens), one staminodal (3 staminodes), filaments covered by subulate trichomes (0.3-0.5 mm long); the two outer fertile whorls similar in shape and size, 3.6-4.5 mm long, anther 0.9-1.1 × 0.8-0.9 mm, 4-celled in two rows; the inner whorl with 3 fertile stamens in front of outer tepals, 2.5-3.9 mm long, filament with two globose glands atta-ched ca. 1 mm above the base, 1-1.1 × 0.5-0.6 mm, anther 0.9-1 × 0.4-0.5 mm, 4-celled in two rows; staminodes in front of the internal tepals, 1.7-2.4 mm long, anthers not celled or rarely with four cels in two rows (only seen in a single flower); pistil 3.6-4.5 mm long, 1.3-1.4 mm diameter, strigose from the middle to the apex; style ca. 2.5 mm long; stigma punctiform; fruit a globose berry, 1.3-1.4 cm diame-ter, green at immaturity, red when mature, seed one.
Distribution and habitat:the new species is known to occur in three localities in Boyacá and Santander de-partments (Colombia), in wet forests on slopes and moun-tains terrain, between 450-1300 m elevation in the Ser-ranías de Los Yariguíes and Las Quinchas